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Niles Eldredge

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At the core of punctuated equilibria lies an empirical observation: once evolved, species tend to remain remarkably stable, recognizable entities for millions of years. The observation is by no means new, nearly every paleontologist who reviewed Darwin's Origin of Species pointed to his evasion of this salient feature of the fossil record. But stasis was conveniently dropped as a feature of life's history to he reckoned with in evolutionary biology. And stasis had continued to be ignored until Gould and I showed that such stability is a real aspect of life's history which must be confronted-and that, in fact, it posed no fundamental threat to the basic notion of evolution itself. For that was Darwin's problem: to establish the plausibility of the very idea of evolution, Darwin felt that he had to undermine the older (and ultimately biblically based) doctrine of species fixity. Stasis, to Darwin, was an ugly inconvenience.
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Time Frames: The Rethinking of Darwinian Evolution and the Theory of Punctuated Equilibria, Simon & Schuster: New York NY, 1985, pp.188-189

 
Niles Eldredge

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There can be no question that Darwin had nothing like sufficient evidence to establish his theory of evolution. ... Darwin’s model of evolution ... , being basically a theory of historical reconstruction, ... is impossible to verify by experiment or direct observation as is normal in science ... Moreover, the theory of evolution deals with a series of unique events, the origin of life, the origin of intelligence and so on. Unique events are unrepeatable and cannot be subjected to any sort of experimental investigation. ... His general theory, that all life on earth had originated and evolved by a gradual successive accumulation of fortuitous mutations, is still, as it was in Darwin’s time, a highly speculative hypothesis entirely without direct factual support and very far from that self-evident axiom some of its more aggressive advocates would have us believe. ... One might have expected that a theory of such cardinal importance, a theory that literally changed the world, would have been something more than metaphysics, something more than a myth.

 
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The history of most fossil species includes two features particularly inconsistent with gradualism: 1. Stasis. Most species exhibit no directional change during their tenure on earth. They appear in the fossil record looking much the same as when they disappear; morphological change is usually limited and directionless. 2. Sudden appearance. In any local area, a species does not arise gradually by the steady transformation of its ancestors; it appears all at once and “fully formed.”

 
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